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Proinflammatory cytokines produced by microglia and astrocytes upon activation cause upregulation of MMP-9 expression in endothelial cells, which results in the degradation of tight junction proteins and causes the breakdown of the BBB (223)

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10.1021/bi701559f 95 ShenP.HanL.ChenG.ChengZ.LiuQ

This dual nature of GSH makes it an attractive but challenging target for therapeutic applications