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The folate-cobalamin interaction is pivotal for normal synthesis of purines and pyrimidines and the transfer of the methyl group to cobalamin is essential for the adequate supply of tetrahydrofolate, the substrate for metabolic steps that require folate

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The gamma linkage (between glutamate's side-chain carboxyl group and cysteine's amino group) was unusual and made the molecule resistant to standard peptidases, which explained why it accumulated in cells rather than being rapidly degraded