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The Rac1 activator, phorbol 12-myristate 13-acetate (PMA), significantly increased the protein expression levels of MMP-2, -3, -9, and -19, whereas rhein and the Rac1 inhibitor, NSC23766, produced opposite effects

Flow cytometry was performed using a BD Fortessa instrument (BD) or NovoCyte Quanteon and data were analyzed using FlowJo v10.6.1 software (Tree Star)

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